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Abbreviations 5-oxp: Pyroglutamic acid (5-oxoproline) AMP: Adenosine monophosphate APX: Ascorbate peroxidase ATP: Adenosine triphosphate BEH: Ethylene bridged hybrid CAT: Catalase Chl: Chlorophyll C initial : Initial conductivity C max : Maximum conductivity DAB: 3',3-diaminobenzidine DHAR: Dehydroascorbate reductase DW: Distilled water EDTA: ethylenediaminetetraacetic acid EL: Electrolyte leakage F m : Maximum fluorescence F v : Variable fluorescence F v /F m : Photochemical efficiency GR: Glutathione reductase GSH: Glutathione H 2 O 2 : Hydrogen peroxide HPLC-ESI-MS: High-performance liquid chromatography-electrospray ionization-mass spectrometer m/z: Mass-to-charge ratio MDHAR: Monodehydroascorbate reductase NH 3 AC: Ammonium acetate NH 3 OH: Ammonia PBS: Phosphate-buffered saline PVPP: polyvinylpyrrolidone TCA: Tricarboxylic acid UDP-glucose: Uridine diphosphate-glucose (UDP-glucose) UGPase: Uridine diphosphate-glucose pyrophosphorylase UMP: Uridine monophosphate References Akram M (2014) Citric acid cycle and role of its intermediates in metabolism